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Adjusting the light levels may not be enough to even out all the darks and lights in a picture. To further correct the picture, you can adjust the shadows and highlights.
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Adjusting midtone contrast will increase or decrease the difference between darks and lights in your photo. You can see in the first image that there is a greater difference between the two than before adjusting the midtone contrast.
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Darkening highlights is useful for darkening particularly light spots in a picture without having to select that specific area.
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The Lighten Shadows tool is used for lightening particularly dark spots. Spots that contain detail (like the Nikon logo over the lens) may appear solid black; lightening shadows can bring out the detail. Use this tool only in small doses, though, as black objects are usually considered "shadows" in a program, and they will be lightened as well. Overuse of this function can also result in grainy spots.
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